Passive Transport Section 1
Cells live in a liquid environment
where the exchange of substances between the cell and its surroundings must
pass through the cell membrane. We know that there is a significant difference
in the substance content inside and outside the cell, which relates to the
function of the cell membrane. What are the different characteristics of
transmembrane transport for different substances? Water Movement Principle
When a drop of red ink is dropped into
a glass of water, the water quickly turns red, a result of solute molecules
diffusing in water. The rise of the liquid level in the funnel shown in the
"Problem Exploration" is the result of solvent—water molecules from the beaker—diffusing
into the funnel through a semi-permeable membrane. The diffusion of water
molecules (or other solvent molecules) through a semi-permeable membrane is
called osmosis. If there is a concentration difference on both sides of the
semi-permeable membrane, water molecules will osmotically move from the side
with higher relative water content to the side with lower relative water
content. Is the movement of water molecules
through the cell membrane in and out of cells based on the same principle? Does
the cell membrane act as a semi-permeable membrane? Effects of Placing Mammalian Red Blood Cells in
Different Concentrations of Sodium Chloride Solutions
After a period of time, red blood cells
will undergo the following changes. The principles governing water movement in
and out of other animal cells are the same as those governing red blood cells,
both relying on osmosis. Water Movement in Plant Cells
We know that plant cells have a
distinct structure compared to animal cells. Plant cells have a cell wall
outside the cell membrane. Research shows that for water molecules, the cell
wall is fully permeable, allowing water molecules to freely pass through. The
function of the cell wall is mainly to protect and support the cell, with less
elasticity. In mature plant cells, the central vacuole occupies most of the
cell space (Figure 4-2), squeezing the cytoplasm into a thin layer. Thus, the
liquid environment within the cell mainly refers to the cell sap inside the
vacuole. The cell membrane, vacuole membrane, and the cytoplasm between these
two layers are collectively called the protoplast. When discussing water
movement into and out of cells later on, it mainly refers to water passing
through the protoplast into and out of the vacuole. From the above exploration activities,
it can be seen that the protoplast of plant cells acts like a semi-permeable
membrane, and plant cells also undergo absorption and loss of water through
osmosis. When the concentration of cell sap is lower than that of the external
solution, water in the cell sap passes through the protoplast into the external
solution, causing both the cell wall and protoplast to shrink to some extent.
As the cell continues to lose water, due to the greater elasticity of the
protoplast compared to the cell wall, the protoplast gradually separates from
the cell wall, leading to what is known as plasmolysis. When the concentration
of cell sap is higher than that of the external solution, water from the external
solution passes through the protoplast into the cell sap, gradually restoring
the entire protoplast to its original state and causing the gradual recovery of
plasmolysis in plant cells. Substances like water molecules enter
and exit cells through diffusion, without consuming energy released by chemical
reactions inside the cell. This type of transmembrane transport of substances
is called passive transport. Passive transport is further divided into free
diffusion and facilitated diffusion. Free Diffusion and Facilitated Diffusion
Some small molecule substances can
easily pass through the phospholipid bilayer of the cell membrane, such as
oxygen and carbon dioxide. When the concentration of oxygen in the alveoli is
higher than that inside the alveolar cells due to respiratory action, oxygen
diffuses into the alveolar cells. When the concentration of carbon dioxide
inside the cell increases due to respiratory action, carbon dioxide is expelled
from the cell through diffusion into the extracellular fluid. Lipid-soluble
small molecule organic substances such as glycerol, ethanol, and benzene also
pass easily through simple diffusion into and out of cells. Ions and some small molecule organic
substances like glucose and amino acids cannot freely pass through the cell
membrane. Special proteins embedded in the membrane assist these substances in
crossing the membrane along the concentration gradient; these proteins are
called transport proteins. This diffusion of substances into and out of cells
facilitated by transport proteins is called facilitated diffusion (Figure 4-4),
also known as facilitated diffusion. Transport proteins can be divided into
carrier proteins and channel proteins. Carrier proteins only allow molecules or
ions that fit their binding sites to pass through, and their conformation
changes with each transport event; channel proteins only allow molecules or
ions of appropriate size and charge to pass through their channel, without
requiring direct binding to the protein (Figure 4-5). In the past, it was generally believed
that water molecules entered and exited cells through free diffusion, but later
research showed that water molecules predominantly use aquaporins on the cell
membrane to facilitate diffusion into and out of cells. Since both free diffusion and
facilitated diffusion transport substances across the membrane along the
concentration gradient without consuming energy produced by chemical reactions
inside the cell, the size of the concentration gradient inside and outside the
membrane directly affects the rate of substance transport. However, facilitated
diffusion requires transport proteins, so the rate of transport for certain
substances also depends on the quantity of transport proteins. |
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